"makulo" meaning in Ido

{
  "derived": [
    {
      "_dis1": "0 0",
      "english": "spot",
      "word": "makuleto"
    },
    {
      "_dis1": "0 0",
      "english": "to stain, blotch, blot",
      "word": "makulizar"
    }
  ],
  "etymology_templates": [
    {
      "args": {
        "1": "io",
        "2": "eo",
        "3": "makulo"
      },
      "expansion": "Esperanto makulo",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "en",
        "3": "macula"
      },
      "expansion": "English macula",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "fr",
        "3": "macule"
      },
      "expansion": "French macule",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "de",
        "3": "Makel"
      },
      "expansion": "German Makel",
      "name": "bor"
    },
    {
      "args": {
        "1": "macula",
        "2": "macule",
        "3": "Makel",
        "4": "",
        "5": "",
        "6": "",
        "eo": "makulo"
      },
      "expansion": "Borrowed from Esperanto makulo, English macula, French macule, German Makel.",
      "name": "io-bor"
    }
  ],
  "etymology_text": "Borrowed from Esperanto makulo, English macula, French macule, German Makel.",
  "forms": [
    {
      "form": "makuli",
      "tags": [
        "plural"
      ]
    }
  ],
  "head_templates": [
    {
      "args": {
        "1": "io",
        "2": "noun",
        "head": ""
      },
      "expansion": "makulo",
      "name": "head"
    },
    {
      "args": {
        "1": "makul"
      },
      "expansion": "makulo (plural makuli)",
      "name": "io-noun"
    }
  ],
  "lang": "Ido",
  "lang_code": "io",
  "pos": "noun",
  "senses": [
    {
      "glosses": [
        "spot (usually), sometimes blot or stain"
      ],
      "id": "en-makulo-io-noun-nK7b2jlA",
      "links": [
        [
          "spot",
          "spot"
        ],
        [
          "blot",
          "blot"
        ],
        [
          "stain",
          "stain"
        ]
      ]
    },
    {
      "categories": [
        {
          "kind": "topical",
          "langcode": "io",
          "name": "Anatomy",
          "orig": "io:Anatomy",
          "parents": [
            "Biology",
            "Medicine",
            "Sciences",
            "All topics",
            "Fundamental"
          ],
          "source": "w"
        },
        {
          "_dis": "29 71",
          "kind": "other",
          "name": "Ido entries with incorrect language header",
          "parents": [
            "Entries with incorrect language header",
            "Entry maintenance"
          ],
          "source": "w+disamb"
        }
      ],
      "glosses": [
        "macula"
      ],
      "id": "en-makulo-io-noun-2UTdTWo8",
      "links": [
        [
          "anatomy",
          "anatomy"
        ],
        [
          "macula",
          "macula"
        ]
      ],
      "raw_glosses": [
        "(anatomy) macula"
      ],
      "topics": [
        "anatomy",
        "medicine",
        "sciences"
      ]
    }
  ],
  "sounds": [
    {
      "ipa": "/maˈkulo/"
    }
  ],
  "word": "makulo"
}

{
  "categories": [
    "Ido entries with incorrect language header",
    "Ido lemmas",
    "Ido nouns",
    "Ido terms borrowed from English",
    "Ido terms borrowed from Esperanto",
    "Ido terms borrowed from French",
    "Ido terms borrowed from German",
    "Ido terms derived from English",
    "Ido terms derived from Esperanto",
    "Ido terms derived from French",
    "Ido terms derived from German",
    "Ido terms with IPA pronunciation"
  ],
  "derived": [
    {
      "english": "spot",
      "word": "makuleto"
    },
    {
      "english": "to stain, blotch, blot",
      "word": "makulizar"
    }
  ],
  "etymology_templates": [
    {
      "args": {
        "1": "io",
        "2": "eo",
        "3": "makulo"
      },
      "expansion": "Esperanto makulo",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "en",
        "3": "macula"
      },
      "expansion": "English macula",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "fr",
        "3": "macule"
      },
      "expansion": "French macule",
      "name": "bor"
    },
    {
      "args": {
        "1": "io",
        "2": "de",
        "3": "Makel"
      },
      "expansion": "German Makel",
      "name": "bor"
    },
    {
      "args": {
        "1": "macula",
        "2": "macule",
        "3": "Makel",
        "4": "",
        "5": "",
        "6": "",
        "eo": "makulo"
      },
      "expansion": "Borrowed from Esperanto makulo, English macula, French macule, German Makel.",
      "name": "io-bor"
    }
  ],
  "etymology_text": "Borrowed from Esperanto makulo, English macula, French macule, German Makel.",
  "forms": [
    {
      "form": "makuli",
      "tags": [
        "plural"
      ]
    }
  ],
  "head_templates": [
    {
      "args": {
        "1": "io",
        "2": "noun",
        "head": ""
      },
      "expansion": "makulo",
      "name": "head"
    },
    {
      "args": {
        "1": "makul"
      },
      "expansion": "makulo (plural makuli)",
      "name": "io-noun"
    }
  ],
  "lang": "Ido",
  "lang_code": "io",
  "pos": "noun",
  "senses": [
    {
      "glosses": [
        "spot (usually), sometimes blot or stain"
      ],
      "links": [
        [
          "spot",
          "spot"
        ],
        [
          "blot",
          "blot"
        ],
        [
          "stain",
          "stain"
        ]
      ]
    },
    {
      "categories": [
        "io:Anatomy"
      ],
      "glosses": [
        "macula"
      ],
      "links": [
        [
          "anatomy",
          "anatomy"
        ],
        [
          "macula",
          "macula"
        ]
      ],
      "raw_glosses": [
        "(anatomy) macula"
      ],
      "topics": [
        "anatomy",
        "medicine",
        "sciences"
      ]
    }
  ],
  "sounds": [
    {
      "ipa": "/maˈkulo/"
    }
  ],
  "word": "makulo"
}