"śaδa-" meaning in Proto-Uralic

{
  "descendants": [
    {
      "depth": 1,
      "templates": [],
      "text": "Samoyedic"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Nenets"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "yrk",
            "2": "саˮесь",
            "unc": "1"
          },
          "expansion": ">? Tundra Nenets: саˮесь (saqyesʹ°)",
          "name": "desc"
        }
      ],
      "text": ">? Tundra Nenets: саˮесь (saqyesʹ°)"
    },
    {
      "depth": 1,
      "templates": [],
      "text": "Finno-Ugric:"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Samic"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "se",
            "2": "čuođđit",
            "t": "(of reindeer) to be in rut"
          },
          "expansion": "Northern Sami: čuođđit (“(of reindeer) to be in rut”)",
          "name": "desc"
        }
      ],
      "text": "Northern Sami: čuođđit (“(of reindeer) to be in rut”)"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Ugric"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "hu",
            "2": "szalad"
          },
          "expansion": "Hungarian: szalad",
          "name": "desc"
        }
      ],
      "text": "Hungarian: szalad"
    }
  ],
  "head_templates": [
    {
      "args": {
        "1": "urj-pro",
        "2": "verb"
      },
      "expansion": "*śaδa-",
      "name": "head"
    }
  ],
  "lang": "Proto-Uralic",
  "lang_code": "urj-pro",
  "original_title": "Reconstruction:Proto-Uralic/śaδa-",
  "pos": "verb",
  "senses": [
    {
      "glosses": [
        "to run"
      ],
      "id": "en-śaδa--urj-pro-verb-o9eMoC-L",
      "links": [
        [
          "run",
          "run"
        ]
      ],
      "tags": [
        "reconstruction"
      ]
    },
    {
      "categories": [
        {
          "_dis": "34 66",
          "kind": "other",
          "name": "Pages with 1 entry",
          "parents": [],
          "source": "w+disamb"
        },
        {
          "_dis": "23 77",
          "kind": "other",
          "name": "Pages with entries",
          "parents": [],
          "source": "w+disamb"
        },
        {
          "_dis": "15 85",
          "kind": "other",
          "name": "Proto-Uralic entries with incorrect language header",
          "parents": [
            "Entries with incorrect language header",
            "Entry maintenance"
          ],
          "source": "w+disamb"
        }
      ],
      "glosses": [
        "to be in rut, to be in heat"
      ],
      "id": "en-śaδa--urj-pro-verb-Ju1H8btQ",
      "links": [
        [
          "rut",
          "rut"
        ],
        [
          "heat",
          "heat"
        ]
      ],
      "raw_glosses": [
        "(of animals) to be in rut, to be in heat"
      ],
      "raw_tags": [
        "of animals"
      ],
      "tags": [
        "reconstruction"
      ]
    }
  ],
  "word": "śaδa-"
}

{
  "categories": [
    "Pages with 1 entry",
    "Pages with entries",
    "Proto-Uralic entries with incorrect language header",
    "Proto-Uralic lemmas",
    "Proto-Uralic verbs"
  ],
  "descendants": [
    {
      "depth": 1,
      "templates": [],
      "text": "Samoyedic"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Nenets"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "yrk",
            "2": "саˮесь",
            "unc": "1"
          },
          "expansion": ">? Tundra Nenets: саˮесь (saqyesʹ°)",
          "name": "desc"
        }
      ],
      "text": ">? Tundra Nenets: саˮесь (saqyesʹ°)"
    },
    {
      "depth": 1,
      "templates": [],
      "text": "Finno-Ugric:"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Samic"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "se",
            "2": "čuođđit",
            "t": "(of reindeer) to be in rut"
          },
          "expansion": "Northern Sami: čuođđit (“(of reindeer) to be in rut”)",
          "name": "desc"
        }
      ],
      "text": "Northern Sami: čuođđit (“(of reindeer) to be in rut”)"
    },
    {
      "depth": 2,
      "templates": [],
      "text": "Ugric"
    },
    {
      "depth": 3,
      "templates": [
        {
          "args": {
            "1": "hu",
            "2": "szalad"
          },
          "expansion": "Hungarian: szalad",
          "name": "desc"
        }
      ],
      "text": "Hungarian: szalad"
    }
  ],
  "head_templates": [
    {
      "args": {
        "1": "urj-pro",
        "2": "verb"
      },
      "expansion": "*śaδa-",
      "name": "head"
    }
  ],
  "lang": "Proto-Uralic",
  "lang_code": "urj-pro",
  "original_title": "Reconstruction:Proto-Uralic/śaδa-",
  "pos": "verb",
  "senses": [
    {
      "glosses": [
        "to run"
      ],
      "links": [
        [
          "run",
          "run"
        ]
      ],
      "tags": [
        "reconstruction"
      ]
    },
    {
      "glosses": [
        "to be in rut, to be in heat"
      ],
      "links": [
        [
          "rut",
          "rut"
        ],
        [
          "heat",
          "heat"
        ]
      ],
      "raw_glosses": [
        "(of animals) to be in rut, to be in heat"
      ],
      "raw_tags": [
        "of animals"
      ],
      "tags": [
        "reconstruction"
      ]
    }
  ],
  "word": "śaδa-"
}