"inclusive fitness" meaning in English

{
  "etymology_text": "Coined by W. D. Hamilton in his 1964 paper \"The Genetical Evolution of Social Behaviour\".",
  "head_templates": [
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      "args": {
        "1": "?"
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      "expansion": "inclusive fitness",
      "name": "en-noun"
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  ],
  "lang": "English",
  "lang_code": "en",
  "pos": "noun",
  "senses": [
    {
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          "kind": "other",
          "name": "English entries with incorrect language header",
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        {
          "kind": "topical",
          "langcode": "en",
          "name": "Evolutionary theory",
          "orig": "en:Evolutionary theory",
          "parents": [
            "Biology",
            "Sciences",
            "All topics",
            "Fundamental"
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      ],
      "glosses": [
        "the number of offspring equivalents an individual rears, rescues or otherwise supports through its behavior (regardless of who begets them)."
      ],
      "id": "en-inclusive_fitness-en-noun-3fbFIzMa",
      "links": [
        [
          "evolutionary theory",
          "evolutionary theory"
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      ],
      "qualifier": "evolutionary theory",
      "raw_glosses": [
        "(evolutionary theory) the number of offspring equivalents an individual rears, rescues or otherwise supports through its behavior (regardless of who begets them)."
      ],
      "related": [
        {
          "word": "kin selection"
        }
      ],
      "wikipedia": [
        "W. D. Hamilton",
        "inclusive fitness"
      ]
    }
  ],
  "word": "inclusive fitness"
}

{
  "etymology_text": "Coined by W. D. Hamilton in his 1964 paper \"The Genetical Evolution of Social Behaviour\".",
  "head_templates": [
    {
      "args": {
        "1": "?"
      },
      "expansion": "inclusive fitness",
      "name": "en-noun"
    }
  ],
  "lang": "English",
  "lang_code": "en",
  "pos": "noun",
  "related": [
    {
      "word": "kin selection"
    }
  ],
  "senses": [
    {
      "categories": [
        "English countable nouns",
        "English entries with incorrect language header",
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        "English nouns",
        "English nouns with unknown or uncertain plurals",
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        "en:Evolutionary theory"
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      "glosses": [
        "the number of offspring equivalents an individual rears, rescues or otherwise supports through its behavior (regardless of who begets them)."
      ],
      "links": [
        [
          "evolutionary theory",
          "evolutionary theory"
        ]
      ],
      "qualifier": "evolutionary theory",
      "raw_glosses": [
        "(evolutionary theory) the number of offspring equivalents an individual rears, rescues or otherwise supports through its behavior (regardless of who begets them)."
      ],
      "wikipedia": [
        "W. D. Hamilton",
        "inclusive fitness"
      ]
    }
  ],
  "word": "inclusive fitness"
}